If you have four slots that are full, then you have four physical modules. Usually, there is a sticker on the modules or something with a model number. With yours being so old though the stickers could be gone by now. My suspicion is that one of the modules has failed. Try using one module only until you find the bad one. Install each module, one at a time, in the slot that says 1, or DDR1 or something similar. Boot each time and verify that the computer is recognizing the memory. When you get to the module that is bad the computer will probably not boot, and will possibly beep at you. If that happens, remove the module, install the remaining 1GB module back into slot 1 and the other modules into slots 2 and 3. I don't believe the old DDR1 modules on that board performed in dual channel anyhow so it won't matter about the configuration of the 256mb modules.
If you verify a module that is bad you might as well discard it and find or purchase a replacement module. It would be helpful to have the model numbers of the modules in order to tell you if possibly the failure was due to being run at an incompatible voltage or timing. If all of the modules work, then you have a bad slot on the board.