Using 1st/3rd Video slot for SLI?

If i'm using a ASUS M4N82 Deluxe 980a mobo can I put one video card in the 1st PCI-E slot and the 2nd card in the 3rd PCI-E slot and run these cards in SLI with the full x16 speed for both cards? Or can I only run this with the first two PCI-E slots?

http://www.newegg.com/Product/Product.aspx?Item=N82E16813131383
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More about using video slot
  1. Use the 2 blue slots for 2-way SLI. This board also support 3-way Sli (black slot) with supported video cards, GTX & Ultras.
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