Video Card in Second PCI-E 16X Slot

^ NO...the P55 chipsets have limited PCIe lanes...So only about 20 PCIe lanes are allotted to the graphic slots...And in that mobo, when you populated 2 cards, it will operate at x8x8 mode, which is better than other cheaper boards that operate at x16x4 mode when 2 cards are populated...
Only in these P55 boards, can operate 2 cards at x16x16 mode...
http://www.newegg.com/Product/Product.aspx?Item=N82E16813131594
http://www.newegg.com/Product/Product.aspx?Item=N82E16813188058
 

sgt bombulous

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Just to be clear, I only plan to use ONE card, period. My case design is kind of stupid, and the GPU runs into my hard drive bays in the top slot. So, I just want to move it to the 2nd one. If I have to use the top one I still can, but right now I have a rather peculiar rig keeping the 2nd hard drive out of the way.

Thanks again!
 

sgt bombulous

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In addition to the question above, if I were ever to go SLI (say 2 x GTX260's), how much does the x8 limitation effect this? I'm new to this so this is probably going to sound stupid... But if the slot works at 1/2 speed, what is the advantage in adding the 2nd card? I can only imagine it's not quite this simple.