Noctua fan direction for cases with top ventilation?

I just ordered an Azza Genesis 9000W case, and I will be using it with an Asus P8ZZ7-V MB. The case has a removable MB tray where you can rotate the MB 180* so that the PCI slots are at the top and the CPU at the bottom. This "Reverse ATX, R-ATX" mounting style supposedly cools the system better.

The case has 2x140mm intake fans on the bottom, 2x230mm exhaust fans on top, 1x120mm exhaust fan in the rear, and 2x120 optional fans on the right side for the HDD's. I find it odd that using fans on the bottom of the case to blow the air into the 'wall' created by the PCI cards cools better than blowing air between the cards. Anyone know the logic behind this? I'm accustomed to Proliant and System X rackmount servers, and I don't ever recall seeing an airflow design inside those servers where it's blocked by cards.

At any rate, the Noctua would be in between the bottom fans and the PCI cards when mounted in the R-ATX orientation. Since there are exhausts both in the rear and the top, I'm wondering if the Noctua has the flexibility for the fans to blow either rearward or towards the top in both orientations. And if it has that ability, which would be better for this case?

Thanks!